The Geneplug upgrade, unique to Epsilon's Headquarters subfaction, grants them full technology access and a variety of support powers based on their genetic research. Upgrading the Pandora Hub to acquire the complete weapons array and accompanying support powers can give an Epsilon Proselyte the edge he needs to overcome his enemies in battle.
Besides unlocking the full potential of Epsilon Headquarters' genetic and illusory arsenal, as well as their epic unit, the Aerial Fortress Irkalla, the Geneplug is responsible for providing the following support powers:
A sudden cloud of mutation gas can be deployed anywhere in the battlefield to mutate enemy soldiers into loyal Brutes or outright severely damage them. This support power costs $800 and takes 6:30 to recharge.
Behind the scenes
In 3.0 the Geneplug only costed $500 since it only provided Epsilon HQ-exclusive support powers and the Pandora Hub alone fulfilled the role of granting Tier 3. The same goes for the Psychplug and Chemplug.